According to court documents, husband and wife Michael Zottoli and Pamela Mills spied from Seattle in recent years while living in the Decatur Condominiums at the corner of Spring Street and Boren Avenue in Seattle’s First Hill neighborhood before moving to Virginia.

Federal authorities said the two lived in the building while under orders to gather intelligence about U.S. policy through think tanks and government officials.

A university spokesman said Zottoli and Mills both graduated from the University of Washington with business degrees.

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In June 2004, the couple is believed to have boarded a flight from Sea-Tac Airport to Newark N.J., documents said. Two days later, Zottoli was seen meeting co-conspirator Richard Murphy in New York’s Central Park, authorities said.

Three minutes later, Zottoli was carrying a bag with a museum logo on it, the FBI said.

On Monday, Murphy and four accused accomplices appeared in a federal courtroom in Manhattan. Zottoli and Mills appeared in a courtroom in Virginia.

All suspected spies are being held without bail. An 11th defendant, a man accused of delivering money to the agents, remains at large.

Court documents said Mills is a Canadian who has lived in the U.S. since 2003.

Zottoli said he was born in the U.S., but the FBI said he has lived in the country only since 2001.

The series of arrests of purported deep cover agents followed a multiyear FBI investigation.

Russia's foreign minister said Moscow is waiting for a U.S. explanation about the arrests and two senior lawmakers said some in the U.S. government may be trying to undermine President Barack Obama's warming relations with Russia.

Foreign Minister Sergey Lavrov noted that U.S. authorities announced the arrest days after Russian President Dmitry Medvedev visited the United States.